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===Archaeogenetics=== Mathieson et al. 2018 [[archaeogenetic]] study included three samples from [[Dalmatia]]: two Early & Middle Bronze Age (1631-1521/1618-1513 calBCE) samples from Veliki Vanik (near [[Vrgorac]]) and one Iron Age (805-761 calBCE) sample from Jazinka Cave in [[Krka National Park]]. According to ADMIXTURE analysis they had approximately 60% [[Early European Farmers]], 33% [[Western Steppe Herders]] and 7% [[Western Hunter-Gatherer]]-related ancestry. The male individual from Veliki Vanik carried the [[Human Y-chromosome DNA haplogroup|Y-DNA haplogroup]] [[Haplogroup J-M172|J2b2a1-L283]] while his and two female individuals [[Human mitochondrial DNA haplogroup|mtDNA haplogroup]] were [[Haplogroup I (mtDNA)#I1a1|I1a1]], [[Haplogroup W (mtDNA)|W3a1]] and [[Haplogroup HV (mtDNA)|HV0e]].<ref name="Mathieson2018">{{cite journal | vauthors = Mathieson I, Alpaslan-Roodenberg S, Posth C, Szécsényi-Nagy A, Rohland N, Mallick S, Olalde I, Broomandkhoshbacht N, Candilio F, Cheronet O, Fernandes D, Ferry M, Gamarra B, Fortes GG, Haak W, Harney E, Jones E, Keating D, Krause-Kyora B, Kucukkalipci I, Michel M, Mittnik A, Nägele K, Novak M, Oppenheimer J, Patterson N, Pfrengle S, Sirak K, Stewardson K, Vai S, Alexandrov S, Alt KW, Andreescu R, Antonović D, Ash A, Atanassova N, Bacvarov K, Gusztáv MB, Bocherens H, Bolus M, Boroneanţ A, Boyadzhiev Y, Budnik A, Burmaz J, Chohadzhiev S, Conard NJ, Cottiaux R, Čuka M, Cupillard C, Drucker DG, Elenski N, Francken M, Galabova B, Ganetsovski G, Gély B, Hajdu T, Handzhyiska V, Harvati K, Higham T, Iliev S, Janković I, Karavanić I, Kennett DJ, Komšo D, Kozak A, Labuda D, Lari M, Lazar C, Leppek M, Leshtakov K, Vetro DL, Los D, Lozanov I, Malina M, Martini F, McSweeney K, Meller H, Menđušić M, Mirea P, Moiseyev V, Petrova V, Price TD, Simalcsik A, Sineo L, Šlaus M, Slavchev V, Stanev P, Starović A, Szeniczey T, Talamo S, Teschler-Nicola M, Thevenet C, Valchev I, Valentin F, Vasilyev S, Veljanovska F, Venelinova S, Veselovskaya E, Viola B, Virag C, Zaninović J, Zäuner S, Stockhammer PW, Catalano G, Krauß R, Caramelli D, Zariņa G, Gaydarska B, Lillie M, Nikitin AG, Potekhina I, Papathanasiou A, Borić D, Bonsall C, Krause J, Pinhasi R, Reich D | display-authors = 6 | title = The genomic history of southeastern Europe | journal = Nature | volume = 555 | issue = 7695 | pages = 197–203 | date = March 2018 | pmid = 29466330 | pmc = 6091220 | doi = 10.1038/nature25778 | bibcode = 2018Natur.555..197M }}</ref> Freilich et al. 2021 identify the Veliki Vanik samples as related to the [[Cetina culture]] (EBA-MBA western Balkans). Patterson et al. 2022 study examined 18 samples from the Middle Bronze Age up to Early Iron Age Croatia, which was part of Illyria. Out of the nine Y-DNA samples retrieved, which coincide with the historical territory where Illyrians lived (including tested [[Iapydes]] and [[Liburni]] sites), almost all belonged to the patrilineal line J2b2a1-L283 (>J-PH1602 > J-Y86930 and >J-Z1297 subclades) with the exception of one [[Haplogroup R1b-L2|R1b-L2]]. The mtDNA haplogroups fell under various subclades of H, H1, H3b, H5, J1c2, J1c3, T2a1a, T2b, T2b23, U5a1g, U8b1b1, HV0e. In a three-way admixture model, they approximately had 49-59% EEF, 35-46% Steppe and 2-10% WHG-related ancestry.<ref name="Patterson2022">{{Cite journal|last1=Patterson|first1=Nick|last2=Isakov|first2=Michael|last3=Booth|first3=Thomas|display-authors=1|date=2022|title=Large-scale migration into Britain during the Middle to Late Bronze Age|url=https://www.nature.com/articles/s41586-021-04287-4_reference.pdf|journal=[[Nature (journal)|Nature]]|volume=601|issue=7894|pages=588–594|doi=10.1038/s41586-021-04287-4|pmid=34937049|pmc=8889665 |s2cid=245509501}}</ref> In Lazaridis et al. (2022) key parts of the territory of historical territory of Illyria were tested. In 18 samples from the Cetina culture, all males except for one (R-L51 > Z2118) carried Y-DNA haplogroup J-L283. Many of them could be further identified as J-L283 > Z597 (> J-Y15058 > J-Z38240 > J-PH1602). The majority of individuals carried mtDNA haplogroups J1c1 and H6a1a. The related [[Posušje]] culture yielded the same Y-DNA haplogroup (J-L283 > J-Z38240). The same J-L283 population appears in the MBA-IA Velim Kosa tumuli of Liburni in Croatia (J-PH1602), and similar in LBA-IA Velika Gruda tumuli in Montenegro (J-Z2507 > J-Z1297 > J-Y21878). The oldest J-L283 (> J-Z597) sample in the study was found in MBA [[Shkrel]], northern Albania as early as the 19th century BCE. In northern Albania, IA Çinamak, half of them men carried J-L283 (> J-Z622, J-Y21878) and the other half R-M269 (R-CTS1450, R-PF7563). The oldest sample in Çinamak dates to the first era of post-Yamnaya movements (EBA) and carries R-M269.{{sfn|Lazaridis|Alpaslan-Roodenberg|2022|ps=: Supplementary Files, Table S1 / Supplementary Materials}} Autosomally, Croatian Bronze Age samples from various sites, from Cetina valley and Bezdanjača Cave were "extremely similar in their ancestral makeup",{{sfn|Lazaridis|Alpaslan-Roodenberg|2022|ps=: Supplementary Materials|pp=256}} while from Montenegro's Velika Gruda mainly had an admixture of "Anatolian Neolithic (~50%), Eastern European hunter-gatherer (~12%), and Balkan hunter-gatherer ancestry (~18%)".{{sfn|Lazaridis|Alpaslan-Roodenberg|2022|ps=: Supplementary Materials|pp=220}} The oldest Balkan J-L283 samples have bee found in final Early Bronze Age (ca. 1950 BCE) site of [[Mokrin]] in Serbia and about 100-150 years later in Shkrel, northern Albania.<ref>{{cite journal |last1=Žegarac |first1=Aleksandra |last2=Winkelbach |first2=L |last3=Blöcher |first3=J |last4=Diekmann |first4=Y |last5=Krečković Gavrilović |first5=M |title=Ancient genomes provide insights into family structure and the heredity of social status in the early Bronze Age of southeastern Europe |journal=Scientific Reports |date=2021 |volume=11 |issue=1 |page=10072 |doi=10.1038/s41598-021-89090-x |pmid=33980902 |pmc=8115322 }}</ref><ref name="LazaridisS1Table">{{harvnb|Lazaridis|Alpaslan-Roodenberg|2022|ps=: Supplementary Materials Table S1}}</ref> Aneli et al. 2022 based on samples from EIA Dalmatia argue that the Early Iron Age Illyrians made "part of the same Mediterranean continuum" with the "autochthonous [...] [[Roman Republic]]ans" and had high affinity with [[Daunians]], part of [[Iapygians]] in [[Apulia]], southeastern Italy. Iron Age male samples from Daunian sites have yielded J-M241>J-L283+, R-M269>Z2103+ and I-M223 lineages.<ref>{{cite journal |last1=Aneli |first1=Serena |last2=Saupe |first2=Tina |last3=Montinaro |first3=Francesco |last4=Solnik |first4=Anu |last5=Molinaro |first5=Ludovica |last6=Scaggion |first6=Cinzia |last7=Carrara |first7=Nicola |last8=Raveane |first8=Alessandro |last9=Kivisild |first9=Toomas |last10=Metspalu |first10=Mait |last11=Scheib |first11=Christiana |last12=Pagani |first12=Luca |title=The genetic origin of Daunians and the Pan-Mediterranean southern Italian Iron Age context |journal=Molecular Biology and Evolution |date=2022 |volume=39 |issue=2 |doi=10.1093/molbev/msac014 |pmid=35038748 |pmc=8826970 |url=https://academic.oup.com/mbe/advance-article/doi/10.1093/molbev/msac014/6509524}}</ref> Three Bronze Age males which carry J-L283 have been found in the Late Bronze Age [[Nuragic civilization]] of [[Sardinia]]. This late find in Sardinia in comparison to western Balkan samples suggests a dispersal from the western Balkans towards this region, perhaps via an intermediary group in the Italian peninsula.{{sfn|Lazaridis|Alpaslan-Roodenberg|2022|ps=: Supplementary Materials |p=322}}
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